Corylus plant named ‘OSU 541.147’

ABSTRACT

A new and distinct interspecific hybrid  Corylus  plant named ‘OSU 541.147’ characterized by a vigorous, upright growth habit, the production of nuts with round kernels that fall free of the husk at maturity, and resistance to eastern filbert blight caused by the fungus  Anisogramma anomala  (Peck) E. Müller.

ACKNOWLEDGMENT OF GOVERNMENT SUPPORT

This invention was made with government support under 2016-51181-25412awarded by USDA-National Institute of Food and Agriculture, under58-5358-4-025 awarded by USDA-Agricultural Research Service, and under18-13-202 awarded by USDA-Agricultural Marketing Service. The governmenthas certain rights in the invention.

Botanical denomination: (Corylus americana x Corylus avellana) x Corylusavellana cultivar.

Variety designation: ‘OSU 541.147’.

PARTIES TO JOINT RESEARCH AGREEMENT

Oregon State University, Rutgers, the State University of New Jersey,Board of Regents of the University of Nebraska, and Arbor Day Foundationexecuted a Joint Research Agreement on or before the date subject matterdisclosed and claimed by the present application was made, and suchsubject matter was made as a result of activities undertaken within thescope of the Joint Research Agreement.

BACKGROUND

The present invention relates to a new and distinct cultivar of hybridCorylus plant, botanically known as (Corylus americana x Corylusavellana) x Corylus avellana, and hereinafter referred to by the name‘OSU 541.147’.

The new Corylus plant resulted from a controlled cross of the femaleparent ‘NY 616’ (Corylus americana ‘Rush’ x Corylus avellana‘Barcelona’) (unpatented) x male parent Corylus avellana ‘OSU 226.118’(unpatented) made in February 1990 (FIG. 1). Hybrid seeds resulting fromthe cross were harvested in August 1990. They were provided a period ofmoist chilling, subsequently germinated, and the seedlings were grown inthe greenhouse during the summer of 1991. From this cross, a total of125 seedling trees were planted in a research field in Corvallis, Oreg.,in October 1991. ‘OSU 541.147’ was discovered and selected as a singleplant within that progeny of the stated cross-pollination. It wasoriginally assigned the designation ‘OSU 541.147’, which indicates therow and tree location of the original seedling.

The female parent is a tree labeled ‘NY 616’ in row M and tree 1 (M01)in the variety plot in Corvallis, Oreg. It is an unpatented F₁ Corylusamericana x Corylus avellana selection developed in New York from across of Corylus americana ‘Rush’ x C. avellana ‘Barcelona’ (Slate,1930). ‘Rush’ is a Corylus americana selection from southeasternPennsylvania. ‘Barcelona’ is an old Corylus avellana cultivar from Spainthat is widely distributed in Europe and was introduced in the U.S. inabout 1885 (Mehlenbacher and Miller, 1989). ‘Barcelona’ is more than 200years old and is known under several different names, including‘Castanyera’ in Tarragona (Spain), ‘Grande’ in Asturias (Spain), ‘Gradade Viseu’ in Portugal, and ‘Fertile de Coutard’ in France. The femaleparent contributed incompatibility allele S₂₃ to ‘OSU 541.147’ (Table1). ‘Rush’ carries a dominant allele for eastern filbert blightresistance on linkage group 7 (Bhattarai et al., 2017; Coyne et al.,1998).

TABLE 1 Incompatibility alleles of the NY hybrid hazelnuts developed inGeneva, New York, and the Corylus avellana parents. Also shown are thefield locations at the USDA National Clonal Germplasm Repository. ‘NY616’, the parent of ‘OSU 541.147’, carried S₂₃. Selection S-allelesLocation Listed Parentage ‘Buchanan’ (unpatented) 12 15 N02.53 ‘Rush’ ×‘Barcelona’ ‘Reed’ (unpatented) 12 15 N14.13 ‘Rush’ × ‘Hall's Giant’‘Potomac’ (unpatented)  5 12 N01.53 ‘Rush’ × ‘DuChilly’ ‘NY 104’(unpatented) 14 23 N05.42 ‘Rush’ × ‘DuChilly’ ‘NY 110’ (unpatented) 1012 N02.42 ‘Rush’ × ‘DuChilly’ ‘NY 200’ (unpatented) 15 23 N02.44 ‘Rush’× ‘Hall's Giant’ ‘NY 398’ (unpatented) 15 23 N04.22 ‘Rush’ × ‘RedLambert’ ‘NY 485’ (unpatented) 12 14 N03.28 ‘Rush’ × ‘DuChilly’ ‘NY 529’(unpatented)  3 12 N04.28 ‘Rush’ × ‘Daviana’ ‘NY 588’ (unpatented) 15 23N01.30 ‘Rush’ × ‘Red Lambert’ ‘NY 616’ (unpatented)  1 23 N04.47 ‘Rush’× ‘Barcelona’ ‘NY 1329’ (unpatented)  3 23 N03.24 ‘Rush’ × ‘Cosford’ ‘NY1408’ (unpatented) 11 23 N02.24 ‘Rush’ × ‘Cosford’ ‘NY 1464’(unpatented) 11 23 N01.28 ‘Rush’ × ‘Cosford’ ‘NYF-20’ (unpatented) 14 25N02.22 ‘NY 157’ o.p. ‘NYF-45’ (unpatented) 12 ? N05.47 ‘Snyder’ × ‘NY485’ ‘Barcelona’ (unpatented)  1 2 possibly was ‘Gustav's ‘Cosford’(unpatented)  3 11 Zeller’ ‘DuChilly’ (unpatented) 10 14 syn. ‘ItalianRed’ ‘Hall's Giant’ (unpatented)  5 15 ‘Red Lambert’ unknown(unpatented) ‘White Lambert’  5 10 (unpatented) ‘Gustav's Zeller’ 15 20(unpatented) ‘Rush’ (unpatented) 12 23 by exam of progeny

The male parent ‘OSU 226.118’ is an unreleased selection.

‘OSU 541.147’ was asexually reproduced by rooted suckers in 1997 through2006 in Corvallis, Oreg. The unique features of this new Corylus arestable and reproduced true-to-type in successive generations of asexualreproduction.

SUMMARY

The following traits have been observed and are determined to be theunique characteristics of ‘OSU 541.147’. These characteristics incombination distinguish ‘OSU 541.147’ as a distinct cultivar:

-   -   1. Vigorous and upright plant habit.    -   2. Green to dark-green color of developing and fully expanded        leaves during the spring and summer.    -   3. High level of resistance to eastern filbert blight (EFB)        caused by the fungus Anisogramma anomala (Peck) E. Müller. The        source of this resistance is from C. americana ‘Rush’        (unpatented; Bhattarai, et al., 2017), which differs from the        single dominant allele conferred from ‘Gasaway’ (unpatented,        Mehlenbacher et al., 1991), which protects Corylus avellana        ‘McDonald’ (U.S. Plant Pat. No. 28,200, Mehlenbacher et al.,        2016), ‘Wepster’ (U.S. Plant Pat. No. 27,141, Mehlenbacher et        al., 2014), ‘Dorris’ (U.S. Plant Pat. No. 25,022, Mehlenbacher        et al., 2014), ‘Jefferson’ (unpatented, Mehlenbacher et al.,        2011a), ‘Yamhill’ (unpatented, Mehlenbacher et al., 2009), and        several other Corylus avellana cultivars and pollenizers.    -   4. Expression of incompatibility alleles S₈ and S₂₃ in the        styles.

Comparisons in several replicated plantings in Corvallis, Oreg., CreamRidge, N.J., and East Brunswick, N.J., show that plants of ‘OSU 541.147’differed from plants of the Corylus avellana cultivars ‘Barcelona’(unpatented), ‘Tonda di Giffoni’ (unpatented), ‘Yamhill’, ‘Jefferson’,‘McDonald’, ‘Wepster’ and other cultivars and selections of Corylusavellana known to the Inventors, primarily in their response to EFBpresent in New Jersey, a region where the pathogen is native and highlygenetically diverse (Muehlbauer et al., 2019). They also differed inS-alleles, nut size, kernel percentage (ratio of kernel weight to nutweight), frequency of defects (blank nuts, moldy kernels, twin kernels,etc.), time of pollen shed, and length of the husk or involucre.

For example:

Eastern filbert blight response in New Jersey: In a multi-year trial inEast Brunswick, N.J., ‘OSU 541.147’ showed no eastern filbert blightcompared to the proportion of EFB-diseased wood across the canopycalculated to be 20.4% for ‘Yamhill’, 31.2% for ‘Jefferson’, 48.6% for‘Gasaway’, and 67.0% for ‘Barcelona’.

Pollen shed: ‘OSU 541.147’ generally sheds pollen in East Brunswick,N.J., a week after ‘Ratoli’ (unpatented, minor cultivar from Tarragona,Spain), 2-3 days after ‘Yamhill’ and ‘Santiam’ (unpatented, Mehlenbacheret al., 2007), and 2-3 days prior to ‘Jefferson’ and ‘Gasaway’. InCorvallis, Oreg., OSU 541.147 sheds pollen between ‘McDonald’ (U.S.Plant Pat. No. 28,200) and ‘PollyO’ (U.S. Plant Pat. No. 32,459). ‘OSU541.147’=descriptor-5.

Husk Length: ‘OSU 541.147’ is 1.6 times nut length, slightly shorterthan ‘Barcelona’, while ‘Wepster’ is 2.0 times nut length. Husks haveglandular trichomes.

‘OSU 541.147’ produces small kernels that are suitable for the blanchedkernel market for use in confections and baked goods. ‘OSU 541.147’combines resistance to eastern filbert blight (evaluated againstAnisogramma anomala present in New Jersey, Oregon, Michigan, New York,Pennsylvania, Massachusetts, Wisconsin, and Minnesota [Molnar et al.,2010a]) with round nuts and kernels and moderately good kernelblanching. The tree is vigorous with an upright habit that produces adesirable orchard tree when pruned to a single stem.

Field observations in Corvallis, Oreg., Cream Ridge, N.J., and EastBrunswick, N.J., and results from greenhouse-based inoculationsperformed in New Brunswick, N.J., indicate that ‘OSU 541.147’ expressesresistance to EFB caused by the fungus Anisogramma anomala. Theresistance is conferred by the single dominant allele from Corylusamericana ‘Rush’, found on hazelnut linkage group 7 (Bhattarai et al.,2017), which is unlike the cultivars currently grown in Oregon protectedby the single dominant ‘Gasaway’ resistance allele found on linkagegroup 6 (Mehlenbacher et al., 2006). EFB is now present throughout theWillamette Valley of Oregon where 99% of the U.S. hazelnut crop is grownand is endemic to the eastern U.S. and southern Canada, where it hasseverely limited commercial production of European hazelnut. Fungicideapplications and pruning to remove cankers are currently used to managethe disease in orchards of ‘Barcelona’ and other susceptible cultivarsin the Pacific Northwestern U.S. ‘OSU 541.147’ was selected in theWillamette Valley of Oregon and subsequently evaluated in New Jersey andis adapted to the climate in both regions. ‘OSU 541.147’ is suitable forplanting in areas with high EFB disease pressure. It has shownresistance in the eastern U.S. where the EFB fungus is native andgenetically diverse (Muehlbauer et al., 2019).

The foregoing and other objects and features of the disclosure willbecome more apparent from the following detailed description, whichproceeds with reference to the accompanying figures.

BRIEF DESCRIPTION OF THE DRAWINGS

The accompanying color photographs illustrate the overall appearance ofthe new cultivar, showing the colors as true as it is reasonablypossible to obtain in colored reproductions of this type. Foliage colorsin the photographs may differ slightly from the color values cited inthe detailed botanical description which accurately describe the colorsof the new Corylus.

FIG. 1 shows the pedigree of new cultivar ‘OSU 541.147’.

FIG. 2 shows a tree of the new cultivar ‘OSU 541.147’ hazelnut in Julyof the seventh leaf pruned to a single trunk.

FIG. 3 shows immature nuts and husks of ‘Yamhill’, ‘OSU 541.147’ and‘Barcelona’ in July of the seventh leaf.

FIG. 4 shows mature nuts and husks of ‘OSU 541.147’ growing in NewJersey.

FIG. 5 shows nuts, raw kernels, and blanched kernels of hazelnuts‘Yamhill’, ‘OSU 541.147’ and ‘Barcelona’. Top row nuts, rows 2 and 3 rawkernels, rows 4 and 5 blanched kernels.

FIG. 6 shows the time of female receptivity (bottom, red), pollen shed(top, green), and vegetative budbreak of ‘OSU 541.147’ and otherhazelnut cultivars in Corvallis, Oreg. (December 2018-March 2019).

FIG. 7 shows the time of female receptivity (bottom, red), pollen shed(top, green), and vegetative budbreak of ‘OSU 541.147’ and otherhazelnut cultivars in East Brunswick, N.J. (December 2017 to March2018).

FIG. 8 shows the time of female receptivity (bottom, red), pollen shed(top, green), and vegetative budbreak of ‘OSU 541.147’ and otherhazelnut cultivars in East Brunswick, N.J. (December 2018 to March2019).

SEQUENCE LISTING

The nucleic acid sequences listed in the accompanying sequence listingare shown using standard letter abbreviations for nucleotide bases asdefined in 37 C.F.R. 1.822. Only one strand of each nucleic acidsequence is shown, but the complementary strand is understood asincluded by any reference to the displayed strand. In the accompanyingsequence listing:

SEQ ID NOS: 1-40 are primer sequences that can be used for geneticfingerprinting.

DETAILED DESCRIPTION

The cultivar ‘OSU 541.147’ has not been observed under all possibleenvironmental conditions. The phenotype may vary somewhat withvariations in environment such as temperature and light intensity,without, however, any variance in genotype. The aforementionedphotographs and following observations and measurements describe plantsgrown in East Brunswick, N.J., under commercial practice outdoors in thefield during the spring and summer. The plant used for the photographsand description were from a seven-year-old tree propagated by tie-offlayerage and growing on its own roots. In the following description,color references are made to The Royal Horticultural Society ColourChart, 1966 Edition, except where otherwise noted and where generalterms of ordinary dictionary significance are used. The list of UPOVdescriptors are from the Mar. 28, 1979 Hazelnut guidelines from UPOV.

-   Botanical classification: Corylus cultivar ‘OSU 541.147’.-   Parentage: Female, or seed, parent is ‘NY 616’ (Corylus americana x    Corylus avellana). It was selected from a cross of Corylus americana    ‘Rush’ x Corylus avellana ‘Barcelona’ (Slate, 1930). ‘Rush’ is a    Corylus americana selection from southeastern Pennsylvania.    ‘Barcelona’ is an old Corylus avellana cultivar from Spain that is    widely distributed in Europe and was introduced in the U.S. in about    1885 (Mehlenbacher and Miller, 1989). ‘Barcelona’ is more than 200    years old and is known under several different names, including    ‘Castanyera’ in Tarragona (Spain), ‘Grande’ in Asturias (Spain),    ‘Grada de Viseu’ in Portugal, and ‘Fertile de Coutard’ in France.    ‘Rush’ has been shown to transmit a dominant allele for resistance    to EFB, which has been mapped to a different linkage group than the    allele from ‘Gasaway’. The ‘Rush’ allele that protects ‘OSU 541.147’    has been shown to provide resistance in the presence of multiple    populations of the EFB fungus Anisogramma anomala (Molnar et al.,    2010a, 2019; Bhattarai et al., 2017). Male, or pollen, parent is    Corylus avellana ‘OSU 226.118’ (unpatented breeding selection). ‘OSU    226.118’ is the result of the cross of ‘Tombul Ghiaghli’    (unpatented, Greek origin) x OSU 42.103 (unpatented). ‘OSU 42.103’    is the result of a cross of ‘Montebello’ (unpatented) x ‘Compton’    (unpatented).-   Incompatibility alleles: ‘OSU 541.147’ has incompatibility alleles    S₈ and S₂₃. Hazelnuts are a wind-pollinated, monoecious species that    exhibit a sporophytic self-incompatibility system controlled by a    single locus designated as the S-locus with 33 alleles    (Mehlenbacher, 2014). To develop ‘OSU 541.147’, branches of ‘NY 616’    were emasculated and covered to prevent foreign pollen    contamination. Controlled pollinations used a mixture of pollens of    three breeding selections whose S-alleles were known: ‘OSU 55.129’    (S₂, S₄), ‘OSU 167.002’ (S₃, S₁₀) and ‘OSU 226.118’ (S₁, S₈).    Fluorescence microscopy identified the male parent of ‘OSU 541.147’    (S₈, S₂₃) as ‘OSU 226.118’ because of the common allele S₈. For    comparison, Corylus avellana ‘Sacajawea’ has the alleles S₁ and S₂₂.    ‘Tonda di Giffoni’ has the alleles S₂ and S₂₃, ‘Tonda Pacifica’    (U.S. Plant Pat. No. 22,715, Mehlenbacher et al., 2011b) and    ‘Wepster’ have alleles S₁ and S₂, and ‘McDonald’ has alleles S₂ and    S₁₅ . Corylus americana ‘Rush’ has alleles S₁₂ and S₂₃.-   Propagation (type rooted suckers):    -   -   Time to initiate roots.—About 30 days at 20° C.        -   Time to produce a rooted young plant.—About six months at            22° C.        -   Root description.—Fine to thick; freely branching; creamy            white in color.-   Propagation (type whip grafting):    -   -   Time to budbreak on the scions.—About 14 days at 25° C.        -   Time to produce a grafted plant.—About six months at 25° C.-   Plant description (descriptions are from plants grown in New Jersey    unless otherwise noted):    -   -   General appearance.—Natural habit is perennial shrub, but in            commercial orchards, is a single trunk tree. Upright plant            habit. See FIG. 2.        -   Growth and branching habit.—Freely branching; about 15            lateral branches develop per plant. Pinching, that is,            removal of the terminal apices, enhances branching with            lateral branches potentially forming at every node.        -   Vigor.—Vigorous, upright growth habit.        -   Size.—Plant height is about 3.6 meters; plants grown in            Oregon: plant diameter or spread is about 3.2 meters (in            July, seventh leaf).        -   Trunk at 30 cm above the soil line.—In a trial planted in            Corvallis, Oreg. in 2014, trunk diameter was 7.90 cm and            cross-sectional area was 49 cm² in December, 2019. This is            slightly smaller than ‘Jefferson’ in the same trial.        -   Trunk color.—197B.-   Lateral branch description:    -   -   Length.—About 20.4 cm. Ranges from 14.0 cm to 26.0 cm.        -   Diameter.—About 4.3 mm. Ranges from 3.0 mm to 5.0 mm.        -   Internode length (at base).—About 0.75 cm.        -   Internode length (at tip).—About 3.6 cm. Ranges from 2.5 cm            to 5.0 cm.        -   Texture.—Smooth, pubescent.        -   Strength.—Strong.        -   Color, immature.—143C.        -   Color, mature.—146B.        -   Color of previous seasons branches.—199C.-   Foliage description:    -   -   Arrangement.—Alternate, simple.        -   Length.—About 9.6 cm. Ranges from 7.5 cm to 11.5 cm.        -   Width.—About 7.1 cm. Ranges from 6.0 cm to 9.0 cm.        -   Shape.—Oblong to ovate.        -   Apex.—Obtuse to acute.        -   Base.—Cordate.        -   Margin.—Serrate.        -   Texture, upper and lower surfaces.—Slightly pubescent.        -   Venation pattern.—Pinnate.        -   Color.—Developing foliage — upper surface 144A, lower            surfaces, 144B. Fully expanded foliage — upper surface:            spring and summer, 137B; late summer and fall, 137B. Fully            expanded foliage, lower surface: spring and summer, 137D;            late summer and fall, 137D. Venation, upper surface: spring            and summer, 145A; late summer and fall, 145A.        -   Venation, lower surface.—Spring and summer, 145A; late            summer and fall, 145A.-   Leaf bud description:    -   -   Shape.—Globular.        -   Time of leaf budbreak.—Medium, Descriptor-5. ‘OSU 541.147’            budbreak is about 8 days before ‘Jefferson’, four days after            ‘Yamhill’, and concurrent with ‘Santiam’.        -   Color of leaf buds.—145B.        -   Leaf bud shape (winter).—Ovoid, Descriptor-2.        -   Leaf bud color (winter).—Reddish green, Descriptor-2.-   Petiole description:    -   -   Length.—About 9.3 mm. Ranges from 8.0 cm to 15.0 mm.        -   Diameter.—About 1.8 mm. Ranges from 1.0 mm to 2.0 mm.        -   Texture, upper and lower surfaces.—Pubescent.        -   Color of petiole.—143C.-   Flower description:    -   -   Male inflorescences.—Catkins        -   Color prior to elongation.—176D.        -   Catkin length.—31.7 mm.        -   Female inflorescence style color.—047B.        -   Time of female flowering.—Medium (January 20 to February            10), Descriptor-5.        -   Time of female flowering compared to male            flowering.—Protogyny, Descriptor-1.        -   Involucre constriction.—The involucre is not constricted,            Descriptor-1.        -   Involucre length.—1.6 times length of nut, Descriptor-7.        -   Size of husk indentation.—Medium, Descriptor-7 (similar to            ‘Negret’), about 50% of husk length.        -   Strength of serration of indentation.—Medium, Descriptor-5.        -   Thickness of callus at base.—Thick, Descriptor-7, (similar            to ‘Barcelona’) or about 2 mm.        -   Pubescence on husk.—Present, Descriptor-9.        -   Density of hairiness of involucre.—Strong, Descriptor-7.        -   Jointing of bracts.—On one side, Descriptor-2.-   Nut description:    -   -   Length.—Average 17.4 mm.        -   Width.—Average 17.5 mm.        -   Depth.—Average 14.7 mm.        -   Nut shape.—Globular, Descriptor-2. See FIG. 5.        -   Nut shape index.—(Width+Depth)/2*Length=0.92.        -   Nut compression index.—(Width/Depth)=1.19.        -   Nut weight.—2.62 g (in Corvallis, Oreg.).        -   Kernel weight.—1.16 g (in Corvallis, Oreg.).        -   Kernel percentage (kernel weight/nut weight).—44.1% (in            Corvallis, Oreg.).        -   Number of fruits per cluster.—Three to four. See FIGS. 3 and            4.        -   Nutshell coloration.—165A.        -   Number of stripes on shell.—Medium (about 20), Descriptor-5.        -   Shape of fruit apex.—Flat, Descriptor-1.        -   Prominence of fruit apex.—Slightly prominent, Descriptor-3.        -   Size of fruit pistil scar on shell.—Medium, Descriptor-5.        -   Hairiness of top of fruit.—Medium, Descriptor-5.        -   Curvature of nut basal scar.—Flat, Descriptor-3 (small) and            similar to ‘Negret’. The raised part of the shell has            dimensions 3.3×1.4 mm.        -   Size of basal scar.—Descriptor-5. Average measurements            14.3×13.4 mm.        -   Double kernels.—Absent.        -   Kernel shape.—Globular, Descriptor-2.        -   Shape of kernel in cross-section.—Circular, Descriptor-2.        -   Lateral groove in kernel.—Present.        -   Corkiness of pellicle of kernel.—Slightly corky,            Descriptor-3.        -   Color of the fiber on the kernel.—165A (Royal Horticultural            Society Colour Chart Fifth Edition, 2007).        -   Color of the pellicle under the fiber.—165B (Royal            Horticultural Society Colour Chart Fifth Edition, 2007).-   Disease/pest resistance: Plants of ‘OSU 541.147’ are resistant to    EFB caused by the fungus Anisogramma anomala (Peck) E. Müller.    Plants have not been challenged against all populations of    Anisogramma anomala present in North America (Muehlbauer et al.,    2019); however, ‘OSU 541.147’ has been challenged in field trials    and/or greenhouse inoculations with populations from Oregon, New    Jersey, Minnesota, Michigan, Massachusetts, New York, and    Pennsylvania, and no signs or symptoms of EFB have been observed    (Capik and Molnar, 2012; Molnar et al., 2010a). Resistance is    derived from its grandparent ‘Rush’. ‘OSU 541.147’ has been trialed    in New Jersey since 2002 under high disease pressure with no signs    or symptoms of EFB observed (Capik and Molnar, 2012; Molnar et al.,    2010a, 2019). In Oregon, all trees of ‘OSU 541.147’ have remained    free of EFB. Fungicide applications are not expected to be needed to    control EFB. Susceptibility to bacterial blight caused by    Xanthomonas arboricola pv. corylina has not been quantified, but no    trees in the trials in Oregon and New Jersey were affected.    Susceptibility to bud mite (Phytoptus avellanae Nal.) was rated in    trials in Corvallis, Oreg., in mid-December on a scale of 1 (no    blasted buds) to 5 (many blasted buds). The average rating for ‘OSU    541.147’ was 3.5, indicating moderate susceptibility comparable to    ‘Clark’ and ‘Gamma’ (Table 2). With this rating, chemical control of    mites will occasionally be needed. Buds blasted by bud mites have    not been observed on ‘OSU 541.147’ in New Jersey.

TABLE 2 Performance of 17 genotypes in a replicated trial planted inCorvallis in Spring, 2014. Four trees per selection in a randomizedcomplete block design. Sel Yield (kg/tree) No Selection 2017 2018 2019Total 1 ‘OSU 541.147’ 0.37 2.87 4.58 8.00 2 ‘1250.057’ 1.58 2.70 3.707.98 3 ‘1252.068’ 0.75 3.46 3.93 8.15 4 ‘1253.064’ 0.77 2.40 2.24 5.41 5‘1292.065’ 1.33 2.99 4.83 9.14 6 ‘1300.073’ 1.96 3.33 4.51 9.80 7‘1304.039’ 1.78 2.94 3.62 8.35 8 ‘1307.003’ 1.23 3.30 5.06 9.58 9‘1307.055’ 1.35 3.93 4.06 9.34 10 ‘1308.087’ 1.15 3.70 4.81 9.65 11‘1310.022’ 0.64 2.34 2.16 5.14 12 ‘1339.074’ 1.17 3.89 4.55 9.60 13‘1340.018’ 1.25 4.94 4.53 10.71 14 ‘1341.037’ 1.06 2.08 4.72 7.85 15‘Jefferson’ 2.39 4.77 4.60 11.75 16 ‘McDonald’ 0.78 3.90 5.86 10.53 17‘Wepster’ 2.16 4.59 5.33 12.08 LSD (0.05) 0.57 0.95 1.14 1.81 Sel TrunkTrunk No Selection diameter TCA Yield efficiency Bud mite rating 1 ‘OSU541.147’ 7.89 49.03 0.164 3.50 2 ‘1250.057’ 8.39 55.29 0.143 1.70 3‘1252.068’ 8.42 55.74 0.146 1.80 4 ‘1253.064’ 6.76 35.97 0.151 1.40 5‘1292.065’ 7.79 47.80 0.192 3.10 6 ‘1300.073’ 9.24 67.11 0.146 1.00 7‘1304.039’ 6.82 36.56 0.229 1.10 8 ‘1307.003’ 8.90 62.34 0.152 1.10 9‘1307.055’ 7.71 46.73 0.200 3.10 10 ‘1308.087’ 9.20 66.61 0.144 2.30 11‘1310.022’ 7.97 49.99 0.104 1.00 12 ‘1339.074’ 9.04 64.61 0.149 1.10 13‘1340.018’ 9.48 70.74 0.151 1.10 14 ‘1341.037’ 11.26 99.64 0.079 2.80 15‘Jefferson’ 8.19 52.90 0.222 1.30 16 ‘McDonald’ 9.74 60.07 0.177 2.10 17‘Wepster’ 8.63 58.84 0.210 1.60 LSD (0.05) 0.65 8.78 0.033 1.14 Yieldper tree in each year of evaluation and total of 3 years. There werevery few nuts in 2016 and they were not harvested. Trunk diameter in cm,30 cm above the soil line, calculated from circumference. TCA = Trunkcross-sectional area in cm², calculated from circumference 30 cm abovethe soil line. Yield efficiency is the ratio of total yield per treedivided by trunk cross-sectional area (units kg/cm²). Bud mitesusceptibility rated in mid-December on a scale of 1 (no blasted buds)to 5 (many blasted buds).

-   Temperature tolerance: ‘OSU 541.147’ was selected in Corvallis,    Oreg., and further evaluated in Cream Ridge, N.J. and East    Brunswick, N.J., and is targeted for production in USDA Plant    Hardiness Zones 6a to 7b. Plants of the new Corylus avellana have    been observed to tolerate temperatures from −21 to 38° C.

COMPARATIVE DATA

Disease resistance: ‘OSU 541.147’ differs from existing Corylus avellanacultivars based on its source and type of resistance to eastern filbertblight (EFB) caused by Anisogramma anomala. Commercial cultivarspreviously widely grown in Oregon including ‘Barcelona’ (unpatented),‘Ennis’ (unpatented), ‘Daviana’ (unpatented), ‘Butler’ (unpatented),etc. are highly susceptible to EFB and cannot be grown in the easternU.S. without copious applications of chemical fungicides and heavypruning to remove infected wood. Tree death can occur in the easternU.S. within five years of exposure to the fungus. The more recentlydeveloped cultivars ‘Santiam’ , ‘Yamhill’, ‘Jefferson’, ‘Dorris’,‘Wepster’, and ‘McDonald’ and their associated pollenizers are protectedfrom EFB by a single resistance gene conferred from Corylus avellana‘Gasaway’. This gene provides a high level of resistance in Oregon andWashington where the diversity of the fungus is limited (Muehlbauer etal., 2019), but does not provide a similar level of protection fromdisease in the eastern U.S. where the pathogen is endemic andgenetically diverse (Capik and Molnar, 2012; Molnar et al., 2010b;Muehlbauer et al., 2018). ‘OSU 541.147’ does not carry the single‘Gasaway’ resistance allele. It carries the Corylus americana ‘Rush’allele, which is a different gene on a different chromosome than‘Gasaway’ (Bhattarai, et al., 2017). The allele from ‘Rush’ has beenfound to be very effective against the populations of Anisogrammaanomala present in New Jersey and other locations (Molnar et al., 2010a,2019).

In a multi-year trial in East Brunswick, N.J., completed in winter 2018and spanning more than 8 years of exposure to EFB, the averageproportion of diseased wood (total length of EFB-diseased stems per treedivided by total length of shoot growth) for ‘OSU 541.147’ was 0.0% (noEFB) compared to 20.4% for ‘Yamhill’ (unpatented, Mehlenbacher et al2009), 31.2% for ‘Jefferson’ (unpatented, Mehlenbacher et al. 2011a),and 48.6% for ‘Gasaway’ (unpatented). Previous studies in New Jerseyshowed the proportion of diseased wood of ‘Barcelona’ to be 67.0%,‘Tonda di Giffoni’ 39%, and ‘Sacajawea’ 21% (Capik and Molnar, 2012).

Differences were also observed in the number of cankers and averagecanker length for ‘OSU 541.147’ in comparison to ‘Yamhill’, ‘Jefferson’,and ‘Gasaway’ in the study completed in 2018. ‘OSU 541.147’ expressed nocankers. In contrast, ‘Gasaway’ exhibited an average of 93.0 cankers pertree with an average length of 130.8 cm, ‘Jefferson’ exhibited anaverage of 36.9 cankers per tree with an average length of 72.3 cm, and‘Yamhill’ exhibited an average of 40.5 cankers per tree with an averagelength of 37.9 cm. As reported in Capik and Molnar (2012), and as afurther point of comparison in regard to EFB response, ‘Barcelona’exhibited an average of 20.4 cankers per tree with an average length of61.9 cm, ‘Tonda di Giffoni’ exhibited an average of 39.0 cankers pertree with an average length of 24.5 cm, and ‘Sacajawea’ exhibited anaverage of 7.7 cankers per tree with an average length of 21.5 cm (Capikand Molnar, 2012).

Nut and kernel characteristics. ‘OSU 541.147’ hazelnut is targeted forthe blanched kernel market and specifically for nut production in theeastern United States in USDA Plant Hardiness Zones 6a to 7b where mostexisting cultivars of Corylus avellana cannot be grown due to theimpacts of EFB.

As shown in FIG. 5, the nut shape is round to somewhat oblong. Kernelsare round. The average single nut weight for ‘OSU 541.147’ over threeyears (2017-19) is 2.62 g, average single kernel weight is 1.16 g, withan average kernel percentage of 44.1% (FIG. 4, Table 3). Nut weight inthe same trial was 3.77 g for ‘Jefferson’, 2.67 g for ‘McDonald’ and2.49 g for ‘Wepster’. Kernel weights in the same trial were 1.70 g for‘Jefferson’, 1.39 g for ‘McDonald’ and 1.17 g for ‘Wepster’. Kernelpercentage in the same trial was 45.2% for ‘Jefferson’, 52.3% for‘McDonald’ and 47.0% for ‘Wepster’. ‘OSU 541.147’ nuts and kernels aresignificantly smaller than those of ‘Barcelona’ and ‘Jefferson’, andintermediate between ‘McDonald’ and ‘Wepster’. In a previous trial(Mehlenbacher et al., 2008), ‘Barcelona’ had an average nut weight of3.85 g, average single kernel weight of 1.66 g, and an average kernelpercentage of 43.1%.

In the trial planted in 2014 in Corvallis, Oreg., ‘Barcelona’ had anaverage single nut weight of 3.77 g, an average single kernel weight of1.70, and an average kernel percentage of 45.2%. The trees produced afew nuts in 2016, but were not harvested. Nuts were harvested for threeyears (2017-19), dried, weighed and evaluated. Total nut weight per tree(2017-19) was 8.0 kg for OSU 541.147, which is less than for the checks‘Jefferson’, ‘McDonald’ and ‘Wepster’ (Table 2). Trunk cross-sectionalarea (TCA) was 49.03 cm², or slightly smaller than ‘Jefferson’. Yieldefficiency, the ratio of total yield to TCA, was a respectable 0.164kg/cm² and similar to ‘McDonald’ (0.177 kg/cm²). Trees have an uprightgrowth habit, and vigor similar to C. avellana selections in theCorvallis trials.

Raw kernels of ‘OSU 541.147’ have a medium brown pellicle with a largeamount of attached fiber (average rating was 4.0 on a scale of 1 [nofiber] to 4 [much fiber]; Table 3). Pellicle removal after roasting at150° C. for 15 min and rubbing is rated on a scale of 1 (completepellicle removal) to 7 (no pellicle removal). Slightly less than half ofthe pellicle on ‘OSU 541.147’ kernels is generally removed afterroasting with an average rating of 4.5 (Table 3), a value similar to‘Barcelona’ (4.2 out of 7.0 as described in Mehlenbacher et al., 2008)and ‘Yamhill’ (4.1 out of 7.0 as described in Mehlenbacher et al.2011a). In the trial planted in 2014, pellicle removal after roastingwas better for ‘McDonald’ (3.7 out of 7) and ‘Wepster’ (3.0 out of 7)(Table 3).

The average percentage of good kernels (kernels free of defects) wascalculated for ‘OSU 541.147’ in the trial planted in 2014 and found tobe 85.3% (Table 3). There was on average 9.3% blank nuts, 0.3% moldykernels, 2.4% nuts with shriveled kernels, and 2.5% poorly filled. Thepercentage of good kernels for ‘OSU 541.147’ was considerably higherthan that reported for ‘Barcelona’ in multiple reports from Oregon(60.9% good kernels reported in Mehlenbacher et al. [2008] and 69.4% inMehlenbacher et al. [2013]). The average percentage of good kernels for‘OSU 541.147’ grown in New Jersey is slightly lower than the rangereported in Oregon for ‘Yamhill’, ‘Jefferson’, ‘Dorris’, and ‘McDonald’,however the percentage of moldy nuts was generally higher for thesecultivars. The incidence of twin kernels and moldy kernels with blacktips is less than 0.1% in both OR and NJ.

TABLE 3 Performance of 17 genotypes in a replicated trial planted inCorvallis in Spring, 2014. Four trees per selection in a randomizedcomplete block design. Sel 10-nut 10-kernel Percent No Selection weightweight kernel Fiber Blanching 1 ‘OSU 26.2 11.6 44.1 4.0 4.5 541.147’ 2‘1250.057’ 36.9 16.4 44.5 3.4 4.2 3 ‘1252.068’ 25.7 12.4 48.3 2.6 2.8 4‘1253.064’ 28.4 13.0 45.9 1.9 3.0 5 ‘1292.065’ 28.2 13.1 46.6 3.5 4.3 6‘1300.073’ 25.2 12.9 51.1 2.6 4.2 7 ‘1304.039’ 27.5 12.4 45.1 2.2 3.0 8‘1307.003’ 23.7 11.9 50.2 1.0 3.0 9 ‘1307.055’ 25.7 12.9 50.4 2.2 4.8 10‘1308.087’ 30.2 13.3 44.1 2.9 4.9 11 ‘1310.022’ 28.8 14.0 48.7 3.5 4.912 ‘1339.074’ 30.5 14.6 47.9 2.0 5.8 13 ‘1340.018’ 27.9 13.9 49.7 3.13.3 14 ‘1341.037’ 30.4 13.5 44.3 2.3 2.1 15 ‘Jefferson’ 37.7 17.0 45.22.8 4.1 16 ‘McDonald’ 26.7 13.9 52.3 2.6 3.7 17 ‘Wepster’ 24.9 11.7 47.02.9 3.0 LSD (0.05) 0.6 0.3 0.5 0.2 0.2 Sel Frequency (%) No Selection GDBL BS MO SF PF TW BT 1 ‘OSU 85.3 9.3 0.3 0.3 2.4 2.5 0.1 0.0 541.147’ 2‘1250.057’ 85.4 4.8 0.3 2.0 1.1 6.0 0.5 0.1 3 ‘1252.068’ 83.7 5.9 1.52.1 0.8 4.3 0.5 1.6 4 ‘1253.064’ 84.4 6.6 1.8 3.4 1.8 3.3 0.6 1.0 5‘1292.065’ 83.1 9.8 0.1 1.8 0.3 4.9 0.3 0.2 6 ‘1300.073’ 83.6 3.5 1.83.5 1.6 3.9 0.2 2.3 7 ‘1304.039’ 84.5 4.8 0.8 2.5 1.4 5.4 0.3 0.5 8‘1307.003’ 86.5 7.9 0.3 2.2 1.5 2.1 0.0 0.3 9 ‘1307.055’ 73.6 8.2 0.22.6 1.1 13.0 1.3 0.7 10 ‘1308.087’ 76.7 6.8 0.0 1.8 5.5 5.7 0.1 0.3 11‘1310.022’ 86.9 3.4 0.1 0.5 0.5 1.7 6.0 1.9 12 ‘1339.074’ 81.7 4.9 0.71.4 1.2 10.3 0.3 0.2 13 ‘1340.018’ 73.0 4.8 0.3 2.1 3.4 16.3 0.1 0.2 14‘1341.037’ 88.2 2.6 1.0 2.6 1.8 3.1 0.5 0.5 15 ‘Jefferson’ 76.5 3.8 2.02.2 1.3 13.8 0.6 1.0 16 ‘McDonald’ 86.0 3.8 0.5 2.3 4.0 3.3 0.1 0.0 17‘Wepster’ 80.3 7.6 0.2 1.3 0.7 8.7 0.0 1.4 LSD (0.05) 3.0 1.9 0.8 1.21.4 1.7 0.6 0.8 Weight of ten well-filled nuts, and ten kernels ingrams. Percent kernel = (kernel weight/nut weight)‘100, based onwell-filled nuts. Fiber on the kernel pellicle is rated from 1 (none) to4 (heavy fiber). Blanching (pellicle removal) is rated from 1 (completepellicle removal) to 7 (no pellicle removal) after roasting at 275° F.for 15-20 minutes and rubbing. Frequency of good nuts and 7 types ofdefects is from two 50-nut samples per tree, averaged over four treesand three years of observation. GD = good kernels, BL = blanks, BS =brown stain, MO = moldy kernels, SH = shriveled kernels, PF = poorlyfilled nuts, TW = twins, and BT = kernels with black tips.

Nut maturity date. The nuts of ‘OSU 541.147’ are typically borne inclusters of 3-4 in husks about 60% longer than the nuts. The husks openas they dry at maturity. About 85% of the nuts fall free of the husk atmaturity (range 75-90%). The other 15% of the nuts come out of the husksas they move through the harvester. When mature, the shells are mediumbrown in color (165A). Harvest date on average is a few days later than‘Jefferson’ when grown in East Brunswick, N.J., and Corvallis, Oreg.(Table 4).

Incompatibility and pollinizers. The trees set a moderate to high numberof catkins that shed pollen in early season 2-3 days after to ‘Yamhill’.Pollen has been collected and germinated on agar medium and bothquantity and viability appear to be good. ‘OSU 541.147’ hasincompatibility alleles S₈ and S₂₃ as determined by fluorescencemicroscopy. Both alleles are expressed in the female flowers but only S₈is expressed in the pollen due to dominance. By convention, allelesexpressed in the pollen are underlined.

TABLE 4 Harvest dates (2018 and 2019) in the replicated trial planted inCorvallis in 2014, and estimated percentage of nuts on the ground (vs.in the tree) on that date. Also shown for the nuts on the ground is anestimated percentage of nuts free of the husk. 2018 Harvest 2019 Harvest% % % % Selection Date down free Selection Date down free ‘05U541.147’10/5 76 80 ‘541.147’ 10/7 82 88 ‘1252.068’ 10/4 98 98 ‘1252.068’ 10/3 9391 ‘1341.037’ 9/28 93 98 ‘1341.037’ 9/23 99 97 ‘Barcelona’ 10/8 97 96‘Barcelona’ 10/8 99 94 ‘Jefferson’ 10/5 90 83 ‘Jefferson’ 10/7 94 84‘McDonald’ 9/20 92 97 ‘McDonald’ 9/23 99 97 ‘Wepster’ 10/5 99 96‘Wepster’ 9/23 96 76

In Corvallis, Oreg., time of pollen shed and female receptivity wererecorded weekly from early December 2018 to late March 2019 (FIG. 6).Climatic conditions vary each year and impact dates of bloom but notusually the order of progression of bloom among cultivars. In 2019,pollen shed (time of male flowering) of ‘OSU 541.147’ began on January25 and ended on February 25, with peak pollen shed on February 10; thosedates are between those for ‘Wepster’ and ‘PollyO’. The females reachedthe red dot stage on January 15 and remained receptive until February15. Female receptivity spans a shorter time within the receptivity timeof females of ‘Wepster’ and ‘PollyO’. In East Brunswick, N.J., time ofpollen shed and female receptivity were recorded weekly from earlyDecember 2017 to late March 2018 (FIG. 7) and from early December 2018to late March 2019 (FIG. 8). Pollen shed of ‘OSU 541.147’ was about oneweek later than ‘Jefferson’ and ten days later than ‘Yamhill’. Time offemale receptivity overlapped the second half of ‘Yamhill’ and the firsthalf of ‘Jefferson’. Females emerge in early season and are generallyfully receptive around mid-February in New Jersey. Pollinizer cultivarsthat shed compatible pollen in midseason and late midseason arerecommended, with hybrid hazelnut seedlings (Corylus americana x C.avellana) planted as pollenizers in eastern and northern regions wherecold temperatures and fluctuating climatic conditions can affect pollenproduction of C. avellana. In New Jersey, date of leaf budbreak of ‘OSU541.147’ was four days later than ‘Yamhill’ and seven days earlier than‘Jefferson’. In Oregon, date of leaf budbreak of ‘OSU 541.147’ was with‘PollyO’, one day before ‘McDonald’ and four days before ‘Wepster’.

Propagation. Layers of ‘OSU 541.147’ are vigorous and root well, similarto standard cultivars of Corylus avellana.

Additional comparative descriptors. Tables 5 and 6 provide additionaldescriptors distinguishing ‘OSU 541.147’ from various hazelnutvarieties.

TABLE 5 Additional comparative descriptors distinguishing ‘OSU 541.147’from various hazelnut varieties, including measurements and RHS colors(R.H.S. Colour chart, 1966 edition. Royal Horticultural Society (GreatBritain). British Colour Council, London). ‘OSU Character Unit 541.147’‘PollyO’ ‘McDonald’ Lateral branch length cm 20.4 45.6 33.5 (terminalshoot) Lateral branch diameter mm 4.3 4.4 5.0 (terminal shoot) Internodelength cm 3.6 3.4 3.2 (above base) Branch texture smooth, smooth.smooth, glabrous glabrous glabrous Branch strength strong strong strongBranch color, immature RHS 143C 139D 139D Branch color, mature RHS 146B177D 177D Leaf length cm 9.6 13.3 10.4 Leaf width cm 7.1 11.8 8.7 Leafshape oblong to oblong to oblong to ovate ovate ovate Leaf apex obtuseto obtuse to obtuse to acute acute acute Leaf base cordate cordatecordate Leaf margin serrate senate serrate Leaf texture slightlyslightly slightly pubescent pubescent pubescent Leaf venation patternpinnate pinnate pinnate Leaf color developing RHS 144 A 141C 141C leafupper Leaf color developing RHS 144B 139C 139C leaf lower Leaf colorfully expanded leaf RHS 137B 146 A 139B upper, Spr Sum Leaf color fullyexpanded leaf RHS 137B 146 A 139B upper, ea Fall Leaf color fullyexpanded leaf RHS 137D 146B 139C lower, Spr Sum Leaf color fullyexpanded leaf RHS 137D 146B 139C lower, ea Fall Leaf veins, upper. RHS145 A 146 A 139C Spring Summer Leaf veins, upper, ea Fall RHS 145 A 146A 139C Leaf veins, lower. RHS 145 A 146D 139D Spring Summer Leaf veins,lower, ea Fall RHS 145 A 146D 139D Petiole length mm 9.3 16.3 27 Petiolediameter mm 1.8 1.6 1.8 Petiole texture, upper & pubescent pubescentpubescent lower surfaces Petiole color, upper. RHS 143C 146C 139D SpringSummer Petiole color, upper, ea Fall RHS 143C 146C 139D Petiole color,lower, RHS 143C 146D 139D Spring Summer Petiole color, lower, ea FallRHS 143C 146D 139D Catkin color prior RHS ND 138B 176D to elongationFemale flower style color RHS 047B 047B 047B Nut length mm 19.9 19.518.4 Nut width mm 18.8 19.8 18.9 Nut depth mm 15.6 17.2 17.9 Nut shaperound round round Nut shape index [(W + D)/2L] ratio 0.86 0.95 1.00 Nutcompression index (W/D) ratio 1.21 1.15 1.05 Nut shell color RHS 165A164 A 164 A Nut weight g 2.62 2.88 2.51 Kernel weight g 1.16 1.38 1.29Kernel percentage (KW/NW) % 44.1 47.9 51.5 Character Unit ‘Wepster’‘Felix’ ‘York’ Lateral branch length cm 40 43 38 (terminal shoot)Lateral branch diameter mm 5.4 6.0 6.0 (terminal shoot) Internode lengthcm 3.1 2.8 3.2 (above base) Branch texture smooth, smooth, smooth,glabrous glabrous glabrous Branch strength strong strong strong Branchcolor, immature RHS 139D 152B 152B Branch color, mature RHS 177C 152B199A? Leaf length cm 12.1 11 11 Leaf width cm 10.9 10 10 Leaf shapeoblong to oblong to oblong to ovate ovate ovate Leaf apex obtuse toobtuse to obtuse to acute acute acute Leaf base cordate cordate cordateLeaf margin serrate senate serrate Leaf texture slightly slightlyslightly pubescent pubescent pubescent Leaf venation pattern pinnatepinnate pinnate Leaf color developing RHS 141C 144 A 146B leaf upperLeaf color developing RHS 139C 145 A 146C leaf lower Leaf color fullyexpanded leaf RHS 141B 143 A 146 A upper, Spr Sum Leaf color fullyexpanded leaf RHS 141A 143 A 146 A upper, ea Fall Leaf color fullyexpanded leaf RHS 139C 139C 146C lower, Spr Sum Leaf color fullyexpanded leaf RHS 139C 139C 146C lower, ea Fall Leaf veins, upper. RHS139C 139C 146A Spring Summer Leaf veins, upper, ea Fall RHS 139C 139C146 A Leaf veins, lower. RHS 139D 139D 148D Spring Summer Leaf veins,lower, ea Fall RHS 139D 139D 148D Petiole length mm 27 27 27 Petiolediameter mm 1.8 1.8 1.8 Petiole texture, upper & pubescent pubescentpubescent lower surfaces Petiole color, upper. RHS 139D 139D 139D SpringSummer Petiole color, upper, ea Fall RHS 139D 139D 139D Petiole color,lower, RHS 139D 139D 139D Spring Summer Petiole color, lower, ea FallRHS 139D 139D 139D Catkin color prior RHS 176C 194C shade, 194C toelongation 176D sun Female flower style color RHS 047B 047B 047B Nutlength mm 18.3 18.7 18.0 Nut width mm 19.0 18.9 19.7 Nut depth mm 16.616.7 17.0 Nut shape round round round Nut shape index [(W + D)/2L] ratio0.97 0.95 1.02 Nut compression index (W/D) ratio 1.15 1.13 1.16 Nutshell color RHS 164 A 167 A 164 A Nut weight g 2.39 2.65 2.73 Kernelweight g 1.11 1.32 1.23 Kernel percentage (KW/NW) % 43.9 50.1 46.3 ‘Bur-‘Tonda gundy Character Unit ‘Dorris’ Pacifica’ Lace’ Lateral branchlength cm 51 15 51 (terminal shoot) Lateral branch diameter mm 3.8 5.03.8 (terminal shoot) Internode length cm 3.3 1.3 3.3 (above base) Branchtexture smooth, smooth, smooth, glabrous glabrous glabrous Branchstrength strong strong strong Branch color, immature RHS 152B 178 A 177DBranch color, mature RHS 152B 137 A 177C Leaf length cm 11 12 10.7 Leafwidth cm 10 10 7.8 Leaf shape oblong to oblong to oblong to ovate ovateovate Leaf apex obtuse to obtuse to obtuse to acute acute acute Leafbase cordate cordate cordate Leaf margin serrate serrate serrate Leaftexture slightly slightly slightly pubescent pubescent pubescent Leafvenation pattern pinnate pinnate pinnate Leaf color developing RHS 144 A187 A 187 A leaf upper Leaf color developing RHS 145 A 187 A 183 A leaflower Leaf color fully expanded leaf RHS 143 A 183B 191A upper, Spr SumLeaf color fully expanded leaf RHS 143 A 137 A 191B upper, ea Fall Leafcolor fully expanded leaf RHS 139C 178 A 191B lower, Spr Sum Leaf colorfully expanded leaf RHS 139C 137 A 139B lower, ea Fall Leaf veins,upper. RHS 139C 183B 191A Spring Summer Leaf veins, upper, ea Fall RHS139C 137 A 139C Leaf veins, lower. RHS 139D 178 A 182B Spring SummerLeaf veins, lower, ea Fall RHS 139D 138B 182D Petiole length mm 27 10 27Petiole diameter mm 1.8 2.5 1.8 Petiole texture, upper & pubescentpubescent pubescent lower surfaces Petiole color, upper. RHS 139D 183B191A Spring Summer Petiole color, upper, ea Fall RHS 139D 137 A 139CPetiole color, lower, RHS 139D 178 A 191A Spring Summer Petiole color,lower, ea Fall RHS 139D 138B 139C Catkin color prior RHS 194C shade,176B 177 A to elongation 176D sun Female flower style color RHS 048B183B 183B Nut length mm 19.1 19.0 15.8 Nut width mm 20.7 18.0 15.6 Nutdepth mm 18.2 16.5 12.9 Nut shape round round round Nut shape index[(W + D)/2L] ratio 1.02 0.91 0.85 Nut compression index (W/D) ratio 1.141.09 1.21 Nut shell color RHS 164B 164 A 166C Nut weight g 3.35 2.241.72 Kernel weight g 1.40 1.06 0.76 Kernel percentage (KW/NW) % 43 47 44

TABLE 6 International Union for the Protection of New Varieties ofPlants (UPOV) descriptors distinguishing ‘OSU 541.147’ from varioushazelnut varieties. Character ‘OSU ‘Burgundy No. (UPOV Descriptor)541.147’ ‘PollyO’ Lace’ ‘Dorris’ 1 Plant vigor 5 7 5 4 2 Plant habit 1 55 7 3 Plant shoot density 5 5 5 5 4 Plant suckering 5 4 5 4 5 Shootthickness 5 7 5 7 6 Shoot hairiness 7 3 3 5 7 Shoot lenticels 7 5 3 3 8Leaf bud shape (winter) Ovoid; 2 2 1 3 9 Leaf bud color (winter) Reddish143C,1 3 2 green; 2 10 Time of leaf budburst 6 6 6 6 11 Catkin length(mm) 20.1 20.1; 3 2 8 (winter) mm; 3 12 Catkin color (winter) 138B; 1138B.1 2 2 13 Stigma color 2 047B.1 3 2 14 Time of pollen shed 6 7 6 515 Time of female flowering 6 6 8 6 16 Dichogamy 2 2 3 2 (female vs.male) 17 Leaf blade shape 2 3 2 3 18 Leaf blade size 9.6 × 13.3 × 11.4 ×7 7.1; 3 11.8;5 7.4; 1 19 Leaf blade hairiness 3 3 3 lower side 20Petiole length 9.3 5 7 5 mm; 3 21 Petiole hairiness 7 3 3 5 22 Huskconstriction 1 1 1 1 23 Husk length rcl. to nut 6 6 3 4 length 24 Huskindentation 7 5 5 7 25 Husk serration of 5 5 3 5 indentations 26 Huskthickness of 7 7 3 7 callus at base 27 Husk hairiness 9 9 1 9 (present,absent) 28 Husk hair density 8 3 3 3 29 Husk joining of bracts 1 1 1 230 Nuts per cluster 4 4 2 3 31 Nut size 3 4 2 5 32 Nut shape 3 1 3 1 33Nut shape in cross-section 2 2 1 2 34 Nut shell color 3 2 3 2 35 Nutnumber of stripes 7 3 5 3 on shell 36 Nut shape of top 1 1 4 4 37 Nutapex prominence 7 3 3 3 38 Nut size of pistil scar 3 3 3 3 39 Nuthairiness of top 3 4 5 4 40 Nut size of basal scar 5 3 3 5 41 Nutcurvature 3 2 3 2 of basal scar 42 Double kernels 1 1 1 1 (frequency) 43Kernel size 3 5 2 6 44 Kernel shape 1 1 2 1 43 Kernel cross-section 2 21 2 46 Kernel shape of top 2 2 2 2 47 Kernel shape of base 3 3 2 3 48Kernel lateral groove 1 1 1 1 49 Kernel fiber on skin 9 5 5 3 50 Kernelsize of cavity 5 5 3 7 51 Nut ripening time 7 4 5 6 52 Nut adherence of1 1 1 1 husk after drop 53 Nut percent kernel 3 6 3 3 54 Time of leaffall 3 3 3 3 (November) Character ‘Wep- No. (UPOV Descriptor) ‘York’‘Felix’ ster’ ‘McDonald‘ 1 Plant vigor 5 8 7 5 2 Plant habit 5 3 5 5 3Plant shoot density 5 5 5 5 4 Plant suckering 4 5 3 6 5 Shoot thickness7 7 5 5 6 Shoot hairiness 3 3 5 3 7 Shoot lenticels 5 3 8 Leaf bud shape(winter) 3 2 9 Leaf bud color (winter) 1 2 10 Time of leaf budburst 6 66 6 11 Catkin length (mm) 7 7 5 6 (winter) 12 Catkin color (winter) 2 21 1 13 Stigma color 1 1 1 1 14 Time of pollen shed 6 7 6 5 15 Time offemale flowering 5 5 5 4 16 Dichogamy 2 1 1 1 (female vs. male) 17 Leafblade shape 3 3 3 2 18 Leaf blade size 3 2 3 3 19 Leaf blade hairiness 33 3 3 lower side 20 Petiole length 5 5 5 5 21 Petiole hairiness 5 5 5 522 Husk constriction 1 3 1 1 23 Husk length rel. to 4 5 7 5 nut length24 Husk indentation 7 5 7 7 25 Husk serration of 5 7 7 7 indentations 26Husk thickness of 5 7 5 3 callus at base 27 Husk hairiness 9 9 9 9(present, absent) 28 Husk hair density 3 3 3 3 29 Husk joining of bracts2 2 2 2 30 Nuts per cluster 2 3 3 3 31 Nut size 4 4 3 4 32 Nut shape 1 11 1 33 Nut shape in cross-section 2 2 2 2 34 Nut shell color 2 3 2 2 35Nut number of stripes 5 7 5 5 on shell 36 Nut shape of top 3 1 1 3 37Nut apex prominence 3 5 3 5 38 Nut size of pistil scar 3 3 3 3 39 Nuthairiness of top 3 4 3 4 40 Nut size of basal scar 3 3 3 3 41 Nutcurvature of basal scar 2 2 2 2 42 Double kernels (frequency) 1 I 1 1 43Kernel size 5 5 3 5 44 Kernel shape 1 1 1 1 43 Kernel cross-section 2 22 2 46 Kernel shape of top 2 2 3 1 47 Kernel shape of base 3 3 3 3 48Kernel lateral groove 1 1 1 1 49 Kernel fiber on skin 5 6 5 5 50 Kernelsize of cavity 5 7 5 7 51 Nut ripening time 5 6 5 4 52 Nut adherence ofhusk 1 3 1 1 after drop 53 Nut percent kernel 4 5 4 6 54 Time of leaffall 4 7 4 4 (November)

Microsatellite Marker Analysis: Twenty microsatellite (simple sequencerepeat) markers were used. PCR products were multiplexed post-PCR andsized using capillary electrophoresis (Table 6, and see for exampleBassil et al., Acta Horticulturae 686:105-110, 2005; Gökirmak et al.,Genetic Resources and Crop Evolution 56:147-172, 2009; Gürcan andMehlenbacher. Molecular Breeding 26:551-559, 2010; Gürcan et al.. TreeGenetics and Genomes 6:513-531, 2010).

TABLE 7Primers and annealing temperatures for the microsatellite marker loci used for fingerprinting hazelnut cultivars. Primers  (5'-3')  Lo- Repeat Allele(SEQ ID  (SEQ ID Tm Lo- Refer- Set cus Motif sizes NO:)  NO:) (° C.) nHe Ho PIC r LG cus ence 3 A613 (TC)₁₃₂ 149- Ned- R- 60 14 0.86 0.85 0.850.00 11R A613 Gurcan  (CA)₁ 177 CACACGCCT CCCCTTTCA et al.,  TGTCACTCTCATGTTTGC  2010 TT (1) TT( 2) 2 A614 (TC)₁₇ 125- Hex- R- 60 14 0.85 0.850.84 0.00 6S, A614 Gurcan  (CA)₁₀ 156 TGGCAGAGC GCAGTGGAG 6R et al., NNN(CA)₆ TTTGTCAGC GATTGCTGA 2010 TT (3) CT (4) 3 A616 (AC)₁₁ 136- Fam-R- 60 13 0.85 0.85 0.83 0.00 8R A616 Gurcan  162 CACTCATAC ATGGCTTTTet al.,  CGCAAACTC GCTTCGTTT  2010 CA (5) TG (6) 1 A640 (CT)₁₅₃ 354- F-Fam- 67 11 0.80 0.73 0.77 0.04 10R A640 Gurcan  (CA)₁ 378 TGCCTCTGCCGCCATATA et al.,  AGTTAGTCA ATTGGGATG 2010 TCAAATGTA CTTGTTG  GG (7)(8) 3 B617 (GA)₁₅ 280- Fam- R- 60 9 0.80 0.78 0.78 0.01 8S, B617 Gurcan 298 TCCGTGTTG TGTTTTTGG 8R et al.,  AGTATGGAC TGGAGCGAT  2010 GA (9)G (10) 2 B619 (TC)₂₁ 146- Fam- R- 60 14 0.88 0.88 0.87 0.00 3S, B619Gurcan  180 AGTCGGCTC GCGATCTGA 3R et al.,  CCCTTTTCT CCTCATTTT 2010C(11) TG (12) 4 B634 (AG)₁₅ 218- Hex- R- 60 9 0.76 0.76 0.73 0.00 4RB634 Gurcan  238 CCTGCATCC GTGCAGAGG et al.,  AGGACTCAT TTGCACTCA 2010TA (13) AA (14) 4 B657 (AG)₁₅ 210- Ned- R- 60 8 0.84 0.98 0.82 −0.0811S, B657 Gurcan  228 GAGAGTGCG AGCCTCACC 11R et al.,  TCTTCCTCTTCCAACGAA  2010 GG (15) C (16) 3 B671 (AG)₆NN 221- Hex- R- 60 13 0.860.88 0.84 −0.01 9S, B671 Gurcan  (GA)₁₇ 249 TTGCCAGTG ACCAGCTCT 9Ret al.,  CATACTCTG GGGCTTAAC 2010 ATG (17) AC (18) 2 B709 (GA)₂₁ 219-Ned- R- 60 8 0.74 0.76 0.70 −0.01 5S, B709 Gurcan  233 CCAAGCACGGCGGGTTCT 5R et al.,  AATGAACTC CGTTGTACA 2010 AA (19) CT (20) 1 B733(TC)₁₅ 161- Ned- R- 60 8 0.68 0.68 0.63 0.00 7S, B733 Gurcan  183CACCCTCTT CATCCCCTG 2R et al.,  CACCACCTC TTGGAGTTT 2010 AT (21) TC (22)2 B749 (TC)₁₂ 200- Hex- R- 60 6 0.60 0.64 0.51 −0.03 1R B749 Gurcan  210GGCTGACAA TCGGCTAGG et al.,  CACAGCAGA TGTAGGGTT 2010 AA (23) TT (24) 4B751 (GA)₁₅ 141- Fam- R- 60 7 0.80 0.78 0.77 0.01 7S, B751 Gurcan  153AGCTGGTTC AAACTCAAA 2R et al.,  TTCGACATT TAAAACCCC 2010 CC (25)TGCTC(26) 1 B774 (AG)₁₅ 195- Ned- R- 60 8 0.80 0.80 0.77 0.00 5S, B774Gurcan  213 GTTTTGCGA TGTGTGTGG 5R et al.,  GCTCATTGT TCTGTAGGC 2010CA (27) ACT (28) 3 C115 (TAA)₅ 167- Fam- R- 60 10 0.84 0.90 0.82 0.0354S, C115 Bassil (GAA)₁₂ 225 CATTTTCCG GTTTCCAGA 4R et al., CAGATAATATCTGCCTCC 2005b; CAGG (29) ATATAAT  Gokirmak (30) et al., 2009 3 KG807(TAAA)AA 226- F- Fam- 54 4 0.67 0.78 0.60 −0.07 11 KG807 Gurcan (TAAA)₂A248 AAGCAAGAA CTTACAGAT and (TAAA)₂ AGGGATGGT AAATGGCTC Mehlen- (31)AAA (32) bacher, 2010 1 KG809 (AGG)₆ 333- F- Hex- 55 5 0.66 0.64 0.600.01 4 KG809 Gurcan 345 GGAAGGTGA AGGCATCAG and GAGAAATCA TTCATCCAA Mehlen- AGT (33) (34) bacher, 2010 2 KG811 (GA)₁₇ 240- F- Ned- 58 120.83 0.82 0.81 0.01 2 KG811 Gurcan 278 GAACAACTG AAGGCGGCA and AAGACAGCACTCGCTCAC  Mehlen- AAG (35) (36) bacher, 2010 4 KG827 (CT)₁₃AA 264- Fam-R- 67 9 0.78 0.84 0.75 −0.04 9 KG827 Gurcan (CA)₇ 282 AGAACTCCGGAGGGAGCA and ACTAATAAT AGTCAAAGT Mehlen- CCTAACCCT TGAGAAGAA  bacher,TGC (37) A( 38) 2010 2 KG830 (CT)₁₄ 279- Ned- R- 67 9 0.79 0.78 0.760.00 9 KG830 Gurcan GTATT 311 TGGAGGAAG AAAGCAACT and (CA)₈ TTTTGAATGCATAGCTGA Mehlen- GTAGTAGAG AGTCCAATC  bacher, GA (39) (40)A 2010 Set= Multiplex set, with samples mixed after PCR but before submission forsizing by capillary electrophoresis; Allele size = range of sizes;Primers, forward (F, listed first) and reverse (R, second) (Hex and Famare dyes); Tm = Annealing temperature; n = Number of alleles; He= expected heterozygosity; Ho = observed heterozygosity; PIC= Polymorphism information content; r = frequency of null alleles; LG= linkage group (S is susceptible female parent ‘OSU 252.146’; R isresistant parent ‘OSU 414.062’); Reference is the journal article whereadditional details were published.

Table 8 shows allele sizes at 20 microsatellite markers for ‘OSU541.147’, its female parent ‘NY 616’, and 13 additional cultivars andselections. ‘Rush’, ‘Barcelona’, ‘Montebello’ and ‘Tombul Ghiaghli’ arein the pedigree of ‘OSU 541.147’. Cultivars ‘OSU 541.147’, ‘NY 110’, ‘NY616’, ‘Rush’, ‘Barcelona’ , ‘Montebello’, and ‘Tombul Ghiaghli’ werefingerprinted in 2020. The other cultivars were fingerprinted in 2018for ‘PollyO’ (U.S. Plant Patent Publication No. US-2020-0008334-P1).‘OSU 541.147’ is easily distinguished from all others shown in Table 8.‘OSU 541.147’ shares an allele with its mother ‘NY 616’ at all SSR loci.

TABLE 8 Allele sizes at 20 microsatellite markers ‘OSU ‘NY ‘Monte-‘Tombul Marker 541.147’ 616’ ‘Rush’ ‘Barcelona’ bello’ Ghiaghli’ A613161/161 161/163 153/163 153/161 151/153 153/169 A614 124/150 124/132124/133 124/132 124/132 132/150 A616 144/150 144/150 140/150 144/152152/160 152/160 A640 356/374 356/356 356/356 354/374 362/374 354/374B617 289/289 281/289 281/285 285/289 285/293 293/295 B619 158/170158/172 158/168 158/172 160/166 166/170 B634 228/228 228/240 230/240228/228 232/236 228/232 B657 222/222 214/222 214/214 218/222 218/226210/214 B671 225/249 225/231 231/231 225/229 225/243 229/249 B709229/235 229/235 229/229 227/235 229/235 223/229 B733 173/175 173/175175/175 173/175 175/185 173/175 B749 216/216 208/216 206/208 216/216216/216 216/216 B751 148/150 144/150 150/150 144/154 154/154 148/154B774 205/217 205/213 205/205 209/213 209/213 201/217 C115 182/215174/215 215/215 174/194 174/197 174/182 KG807 252/252 242/252 238/242238/252 238/242 230/253 KG809 339/342 339/339 339/339 339/339 339/348339/342 KG811 263/267 263/267 255/263 261/267 251/267 245/267 KG827278/282 270/282 270/272 282/284 282/282 278/284 KG830 291/295 291/307307/307 291/295 291/295 289/295 Marker ‘NY 110’ ‘Daviana’ ‘Mortarella’‘Negret’ ‘Casina’ A613 153/163 167/179 151/153 153/159 151/153 A614124/132 148/150 132/135 132/132 124/139 A616 140/150 150/152 150/158152/160 144/152 A640 356/374 354/354 354/374 354/368 354/372 B617285/293 289/295 293/295 285/291 285/295 B619 168/168 168/178 158/172166/170 150/158 B634 232/240 228/236 228/236 228/234 228/232 B657214/218 218/226 218/224 218/222 216/218 B671 229/231 239/249 225/243229/237 229/249 B709 229/229 229/229 221/229 223/227 227/233 B733163/175 173/181 175/175 167/175 175/175 B749 208/216 216/216 214/216216/216 214/216 B751 144/150 144/152 144/154 152/154 152/154 B774205/209 213/215 209/220 209/215 209/215 C115 174/215 174/194 182/215182/215 174/197 KG807 242/242 238/252 230/234 238/252 238/252 KG809339/339 339/342 342/342 339/342 339/339 KG811 255/263 255/261 261/267259/267 251/267 KG827 272/284 272/272 282/282 272/284 272/284 KG830295/295 289/295 291/307 295/303 295/303 ‘Tonda Marker ‘Polly0’‘McDonald’ ‘Wepster’ ‘Yamhill’ Pacifica’ A613 153/167 153/169 159/167153/163 159/169 A614 124/158 135/158 135/158 132/158 135/150 A616144/152 150/160 152/160 150/150 150/160 A640 354/354 362/368 368/374354/368 368/374 B617 285/295 293/295 293/295 289/295 293/295 B619158/166 158/172 166/172 158/172 166/172 B634 228/236 222/228 228/228236/236 228/228 B657 218/218 210/218 226/226 218/228 210/226 B671229/249 229/237 239/249 225/243 229/239 B709 223/227 229/229 229/235229/229 229/235 B733 175/181 173/175 173/175 181/185 173/175 B749216/216 214/216 214/216 216/216 214/216 B751 144/154 144/144 144/144152/152 144/154 B774 209/209 209/220 209/213 209/217 209/213 C115194/215 174/197 182/194 197/215 174/182 KG807 238/252 252/252 252/252230/252 228/252 KG809 342/342 339/339 342/342 348/348 339/342 KG811261/267 245/267 257/257 251/261 245/257 KG827 272/284 272/284 270/282282/282 270/284 KG830 291/295 291/295 295/305 291/295 291/295

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We claim:
 1. A new and distinct variety of Corylus plant named ‘OSU541.147’ as illustrated and described.